Home range studies have not been thoroughly conducted among wild populations.
Sometimes seen in pairs (Wild 2000) and ostensibly in family groups (Eaton 2004)
Typically segregate as individuals, in Cameroon (Wild 2000)
Studies of territorial behavior have not been thoroughly conducted among wild populations.
Vocalization (from Dinets 2013 unless otherwise noted)
Olfaction/scent marking (from Weldon and Wheeler 2001)
Aquatic movements (from Frey and Salisbury 2001 unless otherwise noted)