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Komodo Dragon (Varanus komodoensis) Fact Sheet: Bibliography & Resources

Komodo Dragon (Varanus komodoensis)


Ariefiandy A, Purwandana D, Seno A, Ciofi C, Jessop TS. 2013. Can Camera Traps Monitor Komodo Dragons a Large Ectothermic Predator? Gursky-Doyen S, editor. PLoS ONE. 8(3):e58800. doi:10.1371/journal.pone.0058800.

Ariefiandy A, Purwandana D, Natali C, Imansyah MJ, Surahman M, Jessop TS, Ciofi C. 2015. Conservation of Komodo dragons Varanus komodensis in the Wae Wuul nature reserve, Flores, Indonesia: a multidisciplinary approach. International Zoo Yearbook 49: 67-80.

Auffenberg W. 1978. Social and feeding behavior in Varanus komodoensis. In: Greenberg, N. and P. MacLean (Eds.), Behavior and Neurology of Lizards. U.S. Department of Health, Education and Welfare, National Institute of Mental Health, pp. 301-331.

Auffenberg W.  1981. The Behavioral Ecology of the Komodo Monitor.  Gainesville: University of Florida Press, 406 pp.

Auffenberg W.1980. The herpetofauna of Komodo, with notes on adjacent areas. Bulletin of the Florida State Museum Biological Sciences, 25(2):156.

Behler J L. 1988. The Komodo Monitor Consortium-1988. AAZPA, Annual Conference Proceedings,  pp. 159-161.

Birchard GF, T. Walsh, R. Rosscoe, and C.L. Reiben.1995. Oxygen updake by Komodo dragon (Varanus komodoensis) eggs: The energetics of prolonged development in a reptile. Physiological Zoology, 68(4):622-633.

Ciofi,C. 1999. The komodo dragon. Scientific American 280(3):84-91. [Distribution map was obtained online]

Ciofi C. Varanus komodoensis. In: In: Pianka E. and D. King (Eds), Varanoid Lizards of the World, Indiana University Press, Bloomington, Indianapolis. pp.197-204.

Claudino-Sales V. 2019. Komodo National Park, Indonesia. In: Coastal World Heritage Sites (Finkl CW, series ed.). Dordrecht (Netherlands): Springer Nature; p. 551-556.

De Lisle H F. 1996. The Natural History of Monitor Lizards.  Malibar, Florida: Kreiger Publishing Company.

Fitch AJ, Goodman AE, Donnellan SC. 2006. A molecular phylogeny of the Australian monitor lizards (Squamata: Varanidae) inferred from mitochondrial DNA sequences. Australian Journal of Zoology 54:253-269.

Fry, BG, Vidal, N, Norman, IA, Vonk, FI, Scheib, H, Ramjan, SFR, Kuruppu, Fung, SK, Hedges, SB, Richardson, MK, Hodgson, WC, Ignjatovic, V, Summerhayes, R, Kochva, E. 2006. Early evolution of the venom system in lizards and snakes. Nature 439:584-88.

Fry, BG, Wroe, S, Teeuwisse, W, van Osch, MJ, Moreno, K, Ingle, J, McHenry, C, Ferrara, T, Clausen, P, Scheib, H, Winter, KL, Greisman, L, Roelants, K, van der Weerd, L, Clemente, CJ, Giannakis, E, Hodgson, WC, Luz, S, Martelli, P, Krishnasamy, K, Kochva, E, Kwok, HF, Scanlon, D, Karas, J, Citron, DM, Goldstein, EJ, McNaughtan, JE, Norman, JA. 2009. A central role for venom in predation by Varanus komodoensis (Komodo Dragon) and the extinct giant Varanus (Megalania) priscus. Proc. Natl. Acad. Sci. USA. 106: 8969-8974.

Gray C., L.C. Marcus, W.C. McCarten, and T. Sappington. 1967. Amoebiasis in the Komodo dragon, (Varanus komodoensis). International Zoo Yearbook, 6:279-283.

Green B., King D.. 1993.  Goanna; the biology of varanid lizards, New South Wales, Australia:  Wales University Press.
Halverson J and Spelman LH. 2002. Sex determination and its role in management. In: Komodo dragons: biology and conservation, Murphy JB, Ciofi C, de la Panouse C, Walsh T. Zoo and aquarium biology conservation series.

Hildebrandt TB, Goritz F, Pitra C, Spelman L.H, Walsh T, et al. 1996.  Sonomorphological sex determination in subadult Komodo dragons. Proceedings of the Americn Association of Zoolgical Veterinarians, Puerto Vallarta, Mexico. Washington DC: Smithsonian Institution Press; p. 165-176.

Holmes RB, Murray AM, Attia YS, Simons EL, Chatrath P. 2010.

Hudson R, Alberts A., Ellis S, Byers O, (Eds.). 1994.Conservation Assessment and Management Plan for Iguanidae and Varanidae, IUCN/SSC.

Integrated Taxonomic Information System database (ITIS) [Internet]. [accessed 2017 Mar 21]. Varanus komodoensis.

Jones, M.L. 1965. The Komodo Dragon Lizard, Varanus (placovaranus) komodoensis Exhibition and Longevity in Captivity. Unpublished manuscript. 50 pp and Appendix.

King D, King M, Baverstock P. 1991. A new phylogeny of the Varanidae. Mertensiella 2:211-219.

Laver RJ, Purwandan D, Ariefiandy A, Imansyah J, Forsyth D, Ciofi C, Jessop TS. 2012. Life-history and spatial determinants of somatic srowth dynamics in Komodo dragon populations. PLoS 7(9): e45398. doi:10.1371/journal.pone.0045398.

Lutz and Lutz. 1991. Komodo: The Living Dragon, Salem, Oregon: Dimi Press

Miller,P, Castle K. (Eds.) 1995. Komodo Monitor, (Varanus komodoensis) Population and Habitat Viability Assessment (PHVA). Bogor, Indonesia, Briefing Book. IUCN/SSC Conservation Breeding Specialist Group: Apple Valley, MN.

Molnar R. 2004. The long and honorable history of monitors and their kin. In: Pianka E. and D. King (Eds), Varanoid Lizards of the World, Indiana University Press, Bloomington, Indianapolis. pp.10-67.

________ Pianka E. 2004. Biogeography and phylogeny of varanoids. In: Pianka E. and D. King (Eds), Varanoid Lizards of the World, Indiana University Press, Bloomington, Indianapolis. pp.68-76.

Padian K., Olsen PE. 1984. Footprints of the Komodo monitor and the trackways of fossil reptiles. Copeia, 1984(3):862-871. 

Purwandana D, Ariefiandy A, Imansyah MJ, Rudiharto H, Seno A, Ciofi C, Fordham DA, Jessop TS. 2014. Demographic status of Komodo dragons populations in Komodo National Park. Biological Conservation 171: 29-35. doi:10.1016/j.biocon.2014.01.017.

Seal US, Manansang J, Siswomartono D, Suhartono T, Sugarjito J. (Eds.).1995. Komodo Monitor, (Varanus komodoensis), Population and Habitat Viability Assessment Workshop (PHVA). December 4-7, 1995, Taman Safari Indonesian Proceedings Cisuara, Indonesia.

Shaw C. 1963. Dragons big and bold. Oct 1963. ZOONOOZ, Vol. XXXVI (No. 10); p. 3-7.

Spelman L., Cambre RC, Walsh T,. Rosscoe R. 1996. Anesthetic techniques in Komodo dragons. Proceedings of the American Association of Zoolgical Veterinarians, Puerto Vallarta, Mexico.

Sunter G. 2008. Management and reproduction of the Komodo dragon Varanus komodoensis Ouwens 1912 at ZSL London Zoo. International Zoo Yearbook. 42(1):172–182. doi:10.1111/j.1748-1090.2007.00029.x.

Trivedi B. (2003). Trapping Komodo dragons for conservation. National Geographic Today, January 29, 2003. 

UNEP. 2018. Species+: Varanus komodoensis [online database]. Nairobi, Kenya: UNEP [compiled by UNEP-WCMC, Cambridge, UK]. [accessed 2018 Nov 13].

UNESCO. 2019. Komodo National Park [Internet]. [accessed 2019 Apr 10]. Paris: United Nations Educational, Scientific and Cultural Organization.

Walsh T, Rosscoe R, Birchard GF. 1993. Dragon Tales: The history, husbandry and breeding of Komodo monitors at the National Zoological Park. Vivarium, 4(6):23-26.

World Conservation Monitoring Centre. 1996. Varanus komodoensis. The IUCN red list of threatened species [Internet]. Version 2018-1; e.T22884A9396736 [species assessed 1996 Aug 01; page accessed 2018 Nov 13].

Zug GR. 1993. Herpetology. an introductory biology of amphibians and reptiles. Academic Press Inc., San Diego.

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