Because direct observation of a fossil animal's behavior isn't possible, paleontologists use comparison and contrast with living animals for guidance. Tracks can sometimes reveal further clues to behavior. At a fossil site, the mix of plant and animal species gives clues to the ecosystem of that time and place.
Fariña and Blanco (1996)
France et al. (2007)
McDonald & DeIuliis (2008)
McDonald & Pelikan (2005)
Vizcaíno et al. (2001)
Vizcaíno et al. (2008)