Because direct observation of a fossil animal's behavior isn't possible, paleontologists use comparison and contrast with living animals for guidance. Tracks can sometimes reveal further clues to behavior. At a fossil site, the mix of plant and animal species gives clues to the ecosystem of that time and place.
(Van Valkenburgh & Sacco 2002)
Binder et al. (2002)
Hodnett et al. (2009)
Van Valkenburgh et al. (1990)
Van Valkenburgh & Hertel (1993)
Van Valkenburgh & Sacco (2002)