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Guanaco (Lama guanicoe) Fact Sheet: Bibliography & Resources


Baldi RB, Acebes P, Cuellar E, Funes M, Hoces D, Puig S, Franklin WL. 2016. Lama guanicoe. The IUCN red list of threatened species [Internet]. Version 2018.1; eT11186A18540211. [species assessed 2016 Feb 03; page accessed 2018 Nov 10]. doi:10.2305/IUCN.UK.2016-1.RLTS.T11186A18540211.en.

Baldi, Ricardo; Pelliza-Sbriller, Alicia; Elston, David; Albon, Steve (2004) High potential for competition between guanacos and sheep in Patagonia. Journal of Wildlife Management, 68(4): 924-938.

Bank, Michael S.; Sarno, Ronald J.; Franklin, William L. (2003) Spatial distribution of guanaco mating sites in southern Chile: conservation implications. Biological Conservation, 112(3): 427-434.

Bank, Michael S.; Sarno, Ronald J.; Campbell, Nichole K.; Franklin, William L. (2002) Predation of guanacos (Lama guanicoe) by southernmost mountain lions (Puma concolor) during a historically severe winter in Torres del Paine National Park, Chile. Journal of Zoology, 258(2): 215-222.

Beldomenico P M; Uhart M; Bono M F; Marull C; Baldi R; Peralta J L (2003) Internal parasites of free-ranging guanacos from Patagonia. Veterinary Parasitology, 118: 71-77.

Brown, Bruce W. (2000) A review on reproduction in South American camelids. Animal Reproduction Science, 58(3/4): 169-195.

Bulgarella, Mariana; de Lamo, Daniel. (2005) Thermal conductance of guanaco (Lama guanicoe) pelage. Journal of Thermal Biology, 30(8): 569-573.

Cavieres, Lohengrin A.; Fajardo, Alex (2005) Browsing by guanaco (Lama guanicoe) on Nothofagus pumilio forest gaps in Tierra del Fuego, Chile. Forest Ecology and Management, 204(2-3): 237-248.

Cortes, Arturo; Miranda, Eduardo; Rau, Jaime R.; Jimenez, Jaime E. (2003) Feeding habits of guanacos Lama guanicoe in the high Andes of north-central Chile. Acta Theriologica, 48(2): 229-237.

DeGesero, Stephanie. Enrichment options: multifunctional enrichment tool. (2001) Animal Keepers' Forum, 28(6): 247-248.

de Lamo, Daniel A.; Sanborn, Allen F.*; Carrasco, Cynthia D.; Scott, Donald J. (1998) Daily activity and behavioral thermoregulation of the guanaco (Lama guanicoe) in winter. Canadian Journal of Zoology, 76(7): 1388-1393.

Fiori S M; Zalba S M (2003) Potential impacts of petroleum exploration and exploitation on biodiversity in a Patagonian Nature Reserve, Argentina. Biodiversity and Conservation, 12: 1261-1270, 2003.

Fowler, M. E. Husbandry and diseases of camelids. (1996) O I E Revue Scientifique et Technique, 15(1): 155-169. 1996. ("Wildlife husbandry and diseases"; Fowler, M. E. [coord], editor).

Fowler, M.E. (1998) Medicine and Surgery of South American Camelids : llama, alpaca, vicuna, gunaco, 2nd ed. Ames: Iowa State University Press 549p.

Franklin W. (1983) Contrasting socioecologies of South America’s wild camelids: the vicuna and guanaco. In: Eisenberg, J.F. & Kleiman, D.G. Advances in the Study of Mammalian Behavior. Special Publication No. 7. American Society of Mammalogists, Lawarence Kansas : Allen Press, pp 573-629.

Franklin, W. and Here, W. (1990) South American Tylopods In Grzimek’s Encyclopedia of Mammals Vol 5, McGraw-Hill : New York, pp 96-111

Franklin, William L. (1982) Llama language. Llama World 1(2) 6-11.

Franklin, William L. (1983) Contrasting socioecologies of South America’s wild camelids: the vicuna and the guanaco. Am. Soc. Mammal. Spec. Publ.; No. 7. p. 573-631. 11 Mar.

Franklin, William L. (1981) Biology, ecology and relationship to man of the South American camelids. Mmammalian Biology in South America: A Symposium held at the Pymatuning Laboratory of Ecology. May 10-14,. Michael A. Mares and Hugh H. Genoways, editors. p. 457-489.

Franklin, W.L. (1981) Living with guanacos – wild camels of South America. National Geographic, 160(1): pp. 62-75.

Franklin, W. L. (1975) Guanacos in Peru. Oryx, 13(2): p. 191 - 202.

Fraser, M. D. (1998) Diet composition of guanacos (Lama guanicoe) and sheep (Ovis aries) grazing in grassland communities typical of UK uplands. Small Ruminant Research, 29(2): 201-212.

Gonzalez, B.A., Palma, E. Zapata, B and Marin, J.C., (2006) Taxonomic and biogeographical status of guanaco Lama guanicoe (Artiodactyla, Camelidae) Mammal Rev. No 2, 157-178.

Kadwell, Miranda; Fernandez, Matilde; Stanley, Helen F.; Baldi, Ricardo; Wheeler, Jane C.*; Rosadio, Raul; Bruford, Michael W. (2001) Genetic analysis reveals the wild ancestors of the llama and the alpaca. Royal Society of London. Proceedings. Biological Sciences, 268(1485): 2575-2584

Karesh W B; Uhart M M; Dierenfeld E S; Braselton W E; Torres A; House C; Puche H; Cook R A (1998) Health evaluation of free-ranging guanaco (Lama guanicoe). Journal of zoo and wildlife medicine : 29(2):134-41.

Lucherini, M. (1996) Group size, spatial segregation and activity of wild sympatric vicunas Vicugna vicugna and guanacos Lama guanicoe. Small Ruminant Research, May 15, 1996. v. 20 (3), p. 193-198. Amsterdam; New York: Elsevier.

Marin, Juan C.; Spotorno, Angel E.; Gonzalez, Benito A.; Bonacic, Cristian; Wheeler, Jane C.; Casey, Ciara S.; Bruford, Michael W.; Palma, R. Eduardo; Poulin, Elie (2008) Mitochondrial DNA variation and systematics of the guanaco (Lama guanicoe, Artiodactyla: Camelidae) Journal of Mammalogy, 89(2): 269-281.

Marino, Andrea; Baldi, Ricardo (2008) Vigilance Patterns of Territorial Guanacos (Lama guanicoe): The Role of Reproductive Interests and Predation Risk. Ethology, 114(4): pp. 413-423.

Mate M L; Bustamante A; Giovambattista G; de-Lamo D; von-Thuengen J; Zambelli A; Vidal-Rioja L (2005) Genetic diversity and differentiation of guanaco populations from Argentina inferred from microsatellite data. Animal Genetics, 36: 316-321.

Mate, M.L.; Di Rocco, F.; Zambelli, A.; Vidal-Rioja, L. (2004) Mitochondrial DNA structure and organization of the control region of South American camelids. Molecular Ecology Notes, 4 (4): 765-767.

Muller P.L.S. (1776) Erste Classe, Saugende Thiere. In: Des Ritters Carl von Linne vollstandiges Naturalsystem nach der swolften Lateinischen Ausgabe, 1773-1776, pp 1-62 + 3 pls., Suppl. 384 pp, Register, 36 unnumbered p. + 536 pp.

Noss, Andrew J.; Cuellar, Rosa Leny (2001) Community attitudes towards wildlife management in the Bolivian Chaco. Oryx, 35(4): 292-300.

Parreno, V.; Costantini, V.; Cheetham, S.; Blanco Viera, J.; Saif, L. J.; Fernandez, F.; Leoni, L.; Schudel, A.* (2001) First isolation of rotavirus associated with neonatal diarrhoea in guanacos (Lama guanicoe) in the Argentinean Patagonia Region. Journal of Veterinary Medicine. Series B, 48(9): 713-720.

Pugh, D. G. Heat stress in South American camelids. (1996) Proc. Minn. Vet. Med. Assoc. Annu. Conv. 1996: 413-415.

Puig, Silvia; Videla, Fernando; Cona, Monica I.; Roig, Virgilio G. (2008) Habitat use-by guanacos (Lama guanicoe, Camelidae) in northern Patagonia (Mendoza, Argentina) Studies on Neotropical Fauna and Environment, 43(1): 1-9.

Puig S; Videla F; Cona M I (1997) Diet and abundance of the Guanaco (Lama guanicoe Mueller 1776) in four habitats of northern Patagonia, Argentina. Journal of Arid Environments, 36: 343-357.

Puig, Silvia; Videla, Fernando; Monge, Susana; Roig, Virgilio (1996) Seasonal variations in guanaco diet (Lama guanicoe Mueller 1776) and food availability in northern Patagonia, Argentina. Journal of Arid Environments, 34(2): 215-224.

Raedeke KJ. 1979. Population dynamics and socioecology of the guanaco (Lama guanicoe) of Magallanes, Chile. Ph.D. thesis: University of Washington. 424 pp.

Sarno, Ronald J.; Bank, Michael S.; Stern, Hal S.; Franklin, William (2006) Effects of age, sex, season, and social dynamics on juvenile guanaco subordinate behavior. Journal of Mammalogy , 87(1): 41-47.

Sarno, Ronald J.; Bank, Michael S.; Stern, Hal S.; Franklin, William L. (2003) Forced dispersal of juvenile guanacos (Lama guanicoe): Causes, variation, and fates of individuals dispersing at different times. Behavioral Ecology and Sociobiology, 54(1): 22-29.

Sarno, Ronald J.; Clark, William R.; Bank, Michael S.; Prexl, Walter S.; Behl, Michael J.; Johnson, Warren E.; Franklin, William L. (1999) Juvenile guanaco survival: management and conservation implications. Journal of Applied Ecology, 36(6): 937-945.

Sarno, Ronald J.; Franklin, William L. (1999) Maternal expenditure in the polygynous and monomorphic guanaco: suckling behavior, reproductive effort, yearly variation, and influence on juvenile survival. Behavioral Ecology, 10(1): 41-47.

UNEP. 2019. Species+:Lama guanicoe [online database]. Nairobi, Kenya: UNEP [compiled by UNEP-WCMC, Cambridge, UK]. [accessed 2019 Aug 27].

Uhart, Marcela; Baldi, Ricardo (2000) Grim News for Guanacos. Wildlife Conservation, 103(6): p. 12.

Uzal, F. A.; Assis, R. A.; Chang Reissig, E.; Reissig E C (2000) Malignant oedema in a guanaco (Lama guanicoe) Veterinary Record, 147 (12): 336.

Wilson, Paul; William L. Franklin (1985) Male group dynamics and inter-male aggression of guanacos in southern chile. Zeitschrift fur Tierpsychologie, 69(4): 305-328.

Young, JK, and Franklin, WL, (2004) Activity budget patterns in family-group and solitary territorial male guanacos Revista Chilena de Historia Natural 77: 617-625.

Zapata, B.; Fuentes, V.; Bonacic, C.; Gonzalez, B.; Villouta, G.; Bas, F. (2003) Haematological and clinical biochemistry findings in captive juvenile guanacos (Lama guanicoe Mueller 1776) in central Chile. Small Ruminant Research, 48(1); pp. 15-21.

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