77 FR 55530. 2012. Endangered and threatened wildlife and plants; removal of the gray wolf in Wyoming from the Federal List of Endangered and Threatened Wildlife and removal of the Wyoming wolf population's status as an experimental population. Docket No. FWS-R6_E6-2011_0039; FXES11130900000C6–123–FF09E30000. Federal Register. 77(175): 5530-55604.
78 FR 35664. 2013. Endangered and threatened wildlife and plants; removing the gray wolf (Canis lupus) from the List of Endangered and Threatened Wildlife and maintaining protections for the Mexican wolf (Canis lupus baileyi) by listing it as Endangered. Docket No. FWS-HQ-ES-2013-0073; FXES11130900000C2-134-FF09E32000; DOI, Fish and Wildlife Service. Federal Register. 78(114): 35664-35719.
Anderson TM, vonHoldt BM, Candille SI, Musiani M, Greco C, Stahler DR, Smith DW, Padhukasahasram B, Randi E, Leonard JA, Bustamante CD, Ostander EA, Tang H, Wayne RK, Barsh GS. 2009. Molecular and evolutionary history of melanism in North American gray wolves. Science. 323(5919): 1339-1343.
AZA. 2023. Species Survival Statistics [data table report]. Chicago (IL): Lincoln Park Zoo/AZA Population Management Center; [accessed 2023 Apr 18]. https://www.aza.org/species-survival-statistics.
AZA Canid TAG. 2012. Large canid (Canidae) care manual. Association of Zoos and Aquariums. Silver Spring (MD).
Ballard WB, Carby LN, Smith DW. 2003. Wolf interactions with non-prey. In: Mech L D, Boitani L, editors. In: Wolves: behavior, ecology and conservation, Chicago: University of Chicago Press. Pp 259-271.
Bibikov DI. 1982. Wolf ecology and management in the USSR. In: Harrington FH, Paquet PC. Wolves of the world: perspectives of behavior, ecology, and conservation. Park Ridge (NJ): Noyes Publications. Pp. 120-133.
Boitani L. 1982. Wolf management in intensively used areas of Italy. In: Wolves of the world: perspectives of behavior, ecology, and conservation. Park Ridge (NJ): Noyes Publications. Pp. 158-172.
Boitani L, Phillips M, Jhala Y. 2018. Grey wolf Canis lupus. The IUCN red list of threatened species. Version 2018.2; e.T3746A119623865 (species assessed 2018 Aug 30; page accessed 2019 Mar 19). doi:10.2305/IUCN.UK.2018-2.RLTS.T3746A119623865.en.
Brown RW. 1956. Composition of scientific words. Published by the author.
Callan R, Nibbelink NP, Rooney TP, Weidenhoeft JE, Wydeven AP. 2013. Recolonizing wolves trigger a trophic cascade in Wisconsin. Journal of Ecology. 101: 837-845.
Castelló JR. 2018. Canids of the world: wolves, wild dogs, foxes, jackals, coyotes, and their relatives. Princeton (NJ): Princeton University Press.
Comision Nacional de Areas Naturales Protegidas (CONANP). 2009. Programa de Acción para la Conservación de la Especie: Lobo Gris Mexicano (Canis lupus baileyi). CONANP-SEMARNAT. México. 52 pp. http://www.gob.mx/conanp/documentos/programa-de-accion-para-la-conservacion-de-la-especie-lobo-gris-mexicano-canis-lupus-baileyi.
Dorsey GA. 1904. Traditions of the Skidi Pawnee. Boston: for the American Folk-lore Society by Houghton, Mifflin and Co.
Druzhkova AS, Thalmann O, Trifonov VA, Leonard JA, Vorobieva NV, Ovodov ND, Graphodatsky AS, Wayne RK. 2013. Ancient DNA analysis affirms the canid from Altai as a primitive dog. PLoS ONE. 8(3): e57754. doi:10.1371/journal.pone.0057754.
Eisenberg C, Seager ST, Hibbs DE. 2013. Wolf, elk, and aspen food web relationships: context and complexity. Forest Ecology and Management. 299(2013): 70-80.
Fentress JC. 1967. Observations on the behavioral development of a hand-reared male timber wolf. American Zoologist. 7: 339-351.
Ferguson WW 1981. The systematic position of Canis aureus lupaster (Carnivora: Canidae) and the occurrence of Canis lupus in North Africa, Egypt, and Sinai. Mammalia. 45: 459-65.
Fritts, SH, Bangs EE, Fontaine JA, Brewster WG, Gore JF. 1995. Restoring wolves to the northern Rocky Mountains of the United States. In: Carbyn LN, Fritts SH, Seip DR, editors. Ecology and conservation of wolves in a changing world. Edmonton (Alberta): Canadian Circumpolar Institute. Pp. 107-126.
Fuller TK, Mech LD, Cochrane JF. 2003. Wolf population dynamics. In: Mech LD, Boitani L, editors. Wolves: behavior, ecology, and conservation. Chicago: University of Chicago Press. Pp. 161–191.
Geptner VG, Naumov NP, Yurgension PB, Sludskii AA, Chirkova AF, Bannikov AG. 1998. Mammals of the Soviet Union. Vol. 2. Heptner VG, Naumov, editors. Moscow: Vysshaya Shkola Publishers.
Germonpré M, Sablin MV, Stevens RE, Hedges REM, Hofreiter M, Stiller M, Despres VR. 2009. Fossil dogs and wolves from Palaeolithic sites in Belgium, the Ukraine and Russia: osteometry, ancient DNA and stable isotopes. Journal of Archaeological Science. 36: 473–490.
Hall RL, Sharp HS, editors. 1978. Wolf and man: evolution in parallel. New York: Academic Press.
Harrington FH, Asa CS. 2003. Wolf communication. In: Mech LD, Boitani L, editors. Wolves: behavior, ecology and conservation. Chicago: University of Chicago Press. Pp. 66-103.
Harrington FH, Mech LD. 1978. Wolf vocalization. In Hall RL, Sharp HS, editors. Wolf and man: evolution in parallel. New York: Academic Press.
Harrington FH, Mech LD. 1979. Wolf howling and its role in territory maintenance. Behaviour. 68: 207-249.
Harrington FH, Mech LD. 1982. Patterns of homesite attendance in two Minnesota wolf packs. In: Harrington FH, Paquet PC, editors. Wolves of the world: perspectives of behavior, ecology, and conservation. Park Ridge (NJ): Noyes Publications. Pp. 81-105.
Harrington FH, Paquet PC, editors. 1982. Wolves of the world: perspectives of behavior, ecology, and conservation. Park Ridge (NJ): Noyes Publications. P. 118.
Harrison DL. 1968. The mammals of Arabia. London: Ernest Benn.
Hayssen VA, van Tienhoven A, van Tienhoven A. 1993. Asdell's patterns of mammalian reproduction. Ithaca (NY): Comstock Publishing Associates.
Hedrick, P. W., Miller, P. S., Geffen, E. & Wayne, R. 1997. Genetic evaluation of the three captive Mexican wolf lineages. Zool. Biol. 16, 47–69. (doi:10.1002/(SICI)1098-2361(1997)16:1!47::AID-ZOO7O3.0.CO;2-B)
Horowitz A. 2010. Inside of a dog: what dogs see, smell, and know. London: Simon and Schuster.
Integrated Taxonomic Information System database (ITIS). [cited 2014 Mar 01]. Canis lupus (Linnaeus, 1758). Available from: http://www.itis.gov/.
IUCN/SSC. 2019. Wolf — Canis lupus; Large Carnivore Initiative for Europe [Internet]. IUCN/SSC; [accessed 2019 Apr 02]. https://www.lcie.org/Largecarnivores/Wolf.aspx.
Joslin PWB. 1966. Summer activities of two timber wolf (Canis lupus) packs in Algonquin Park [unpublished master's thesis]. Toronto: University of Toronto.
Kelly B, Brown M, Byers O. 2001. Mexican wolf reintroduction program three-year review workshop. Final report. Apple Valley (MN): IUCN/SSC Conservation Breeding Specialist Group.
Kingdon J. 1991. Arabian Mammals: a natural history. London: Academic Press.
Kreeger TJ. 2003. The Internal wolf: physiology, pathology, and pharmacology. In: Mech L D, Boitani L, editors. Wolves: behavior, ecology and conservation. Chicago: University of Chicago Press. Pp 192-217.
Large Carnivore Initiative for Europe 2007. Canis lupus. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.2 [cited 2014 Jan 29]. Available from: www.iucnredlist.org.
Lindblad-Toh K, Wade CM, Mikkelsen TS, Karlsson E, et al. 2005. Genome sequence, comparative analysis and haplotype structure of the domestic dog. Nature. 438(8): 803–819.
Lopez BH. 1978. Of wolves and men. New York: Charles Scribner's Sons.
MacNulty DR, Smith DW, Vucetich JA, Mech LD, Stahler DR, Packer C. 2009. Predatory senescence in aging wolves. Ecology Letters. 12: 1-10.
Mahrenke P, III. 1971. Observations of four wolves killing another wolf. Journal of Mammalogy. 52: 630-631.
McBride RT. 1980. The Mexican wolf (Canis lupus baileyi): a historical review and observations on its status and distribution: progress report to the U.S. Fish and Wildlife Service. Albuquerque (NM): The Service.
McLaren BE, Peterson RO. 1994. Wolves, moose, and tree rings on Isle Royale. Science. 266(5190): 1555-1558.
McNay ME, Stephenson TR, Dale BW. 2006. Diagnosing pregnancy, in utero litter size, and fetal growth with ultrasound in wild, free-ranging wolves. Journal of Mammalogy. 87(1): 85-92.
Mech LD. 1966a. Hunting behavior of timber wolves in Minnesota. Journal of Mammalogy. 47: 347-348.
Mech LD. 1966b. The wolves of Isle Royale. Washington: U.S. US Goverment Printing Office.
Mech LD. 1970. The wolf: the ecology and behavior of an endangered species. Garden City (NY): The Natural History Press.
Mech LD. 1988. Longevity in wild wolves. Journal of Mammalogy. 69(1): 197-198. doi:10.2307/1381776.
Mech LD. 1993. Details of a confrontation between two wild wolves. Canadian Journal of Zoology. 71: 1900-1903.
Mech LD. 1999. Alpha status, dominance, and division of labor in wolf packs. Canadian Journal of Zoology 77:1196-1203.
Mech LD. 2000b. Leadership in wolf, Canis lupus, packs. Canadian Field Naturalist. 114(2): 259-263.
Mech LD. 2002. Breeding season of wolves, Canis lupus, in relation to latitude. Canadian Field Naturalist. 116: 139-140.
Mech LD. 2008. Weight changes in wild wolves, Canis lupus, from ages 2 to 24 months. The Canadian Field-Naturalist. 122: 173-175.
Mech LD. 2012. Is science in danger of sanctifying the wolf? Biological Conservation 150:143-149.
Mech LD, Boitani L, editors. 2003. Wolves: behavior, ecology and conservation. Chicago: University of Chicago Press.
Mech LD, Boitani L. 2004. Grey wolf Canis lupus Linnaeus, 1758 least concern. 2004. In: Sillero-Zubiri C, Hoffmann M, Macdonald DW, editors. Canids: foxes, wolves, jackals and dogs. Status survey and conservation action plan; p. 124-129. IUCN/SSC Canid Specialist Group. Gland, Switzerland: IUCN - The World Conservation Union. https://portals.iucn.org/library/sites/library/files/documents/cover.pdf.
Mech LD, Boitani L (IUCN SSC Wolf Specialist Group). 2010. Canis lupus. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.2. [Cited 2013 Dec 5]. Available from: www.iucnredlist.org.
Mech LD, Cluff HD. 2010. Prolonged intensive dominance behavior between gray wolves, Canis lupus. Canadian Field Naturalist. 124: 215-218.
Mech LD, Korb M. 1978. An unusually long pursuit of a deer by a wolf. Journal of Mammalogy. 59(4): 860-861.
Mech LD, Merrill SB. 1998. Daily departure and return patterns of wolves, Canis lupus, from a den at 80 N latitude. Canadian Field Naturalist. 112(3): 515-517.
Mech LD, Nelson ME. 1990. Non-family wolf, Canis lupus, packs. Canadian Field Naturalist. 104(3): 482-483.
Mech LD, Packard JM. 1990. Possible use of wolf (Canis lupus) den over several centuries. Canadian Field Naturalist. 104: 484-485.
Mech LD, Peterson RO. 2003. Ecosystem effects of wolves. In: Mech LD, Boitani L, editors. Wolves: behavior, ecology and conservation. Chicago: University of Chicago Press. Pp 158-160.
Mech LD, Phillips MK, Smith DW, Kreeger TJ. 1996. Denning behaviour of non-gravid wolves, Canis lupus. Canadian Field Naturalist. 110(2): 343-345.
Mech LD, Adams LG, Meier TJ, Burch JW, Dale BW. 1998. The wolves of Denali. Minneapolis: University of Minnesota Press.
Mech LD, Goyal SM, Paul WJ, Newton WE. 2008. Demographic effects of canine parvovirus on a free-ranging wolf population over 30 years. Journal of Wildlife Diseases. 44: 824-836.
Medjo D, Mech LD. 1976. Reproductive activity in nine and ten month old wolves. Journal of Mammalogy. 57(2): 406-408.
Mendelssohn H. 1982. Wolves in Israel. In: Harrington FH, Paquet PC, editors. Wolves of the world: perspectives of behavior, ecology, and conservation. Park Ridge (NJ): Noyes Publications. Pp. 173-195.
Merkle JA, Krausman PR, Stark DW, Oakleaf JK, Ballard WB. 2009. Summer diet of the Mexican gray wolf (Canis lupus baileyi). Southwestern Naturalist. 54(4): 480-485.
Moulton DG. 1967. Olfaction in mammals. American Zoologist. 7(3): 421-429.
Nowak RM. 1995. Another look at wolf taxonomy. In: Carbyn LN, Fritts SH, Seip DR, editors. Ecology and conservation of wolves in a changing world: Proceedings of the Second North American Symposium on Wolves, Alberta, Canada, 25-27 August 1992. Edmonton(AB): Canadian Circumpolar Institute, University of Alberta. Pp 375-397.
Nowak RM. 1999. Walker's mammals of the world, 6th Edition. Baltimore: The Johns Hopkins University Press.
Nowak RM. 2003. Wolf evolution and taxonomy. In: Mech LD, Boitani L, editors. Wolves: behavior, ecology and conservation. Chicago: University of Chicago Press. Pp 239-258.
NPS website. National Park Service, U.S. Department of the Interior. [cited 2014 Mar 01]. Available from: http://www.nps.gov/yell/naturescience/wolfrest.htm.
Ognev SI. 1962. Mammals of Eastern Europe and northern Asia, Vol. II: Carnivora. Birron A, Cole ZS, translators.
Packard JM. 2003. Wolf behavior: reproductive, social and intelligent. In: Mech LD, Boitani L, editors. Wolves: behavior, ecology and conservation. Chicago: University of Chicago Press. Pp. 35-65.
Pang JF, Kluetsch C, Zou XJ, Zhang AB, Luo LY, et al. 2009. mtDNA data indicate a single origin for dogs south of Yangtze River, less than 16,300 years ago, from numerous wolves. Molecular Biology and Evolution. 26: 2849–2864.
Parker HG, Kim LV, Sutter NB, Carlson S, Lorentzen TD, Malek TB, Johnson GS, DeFrance HB, Ostrander EA, Kruglyak L. 2004. Genetic structure of the purebred domestic dog. Science. 304(5674): 1160-1164.
Parson DR, Nicholopoulos JE. 1995. Status of the Mexican wolf recovery program in the United States. In: Carbyn LN, Fritts SH, Seip DR, editors. Ecology and conservation of wolves in changing world: Proceedings of the Second North American Symposium on Wolves. Edmonton: Canadian Circumpolar Institute, University of Alberta. Pp 141-146.
Peters R, Mech LD. 1978. Scent-marking in wolves. In: Hall RL, Sharp HS, editors. Wolf and man: evolution in parallel. New York: Academic Press.
Range F, Viranyi Z. 2014. Wolves are better imitators of conspecifics than dogs. PloS ONE. 9(1): e86559 DOI: 10.1371/journal.pone.0086559.
Reed JE, Ballard, WB, Gipson PS, Kelly BT, Krausman PR, Wallace MC, Wester DB. 2006. Diet of free-ranging Mexican gray wolves in Arizona and New Mexico. Wildlife Society Bulletin. 34(4): 1127-1133.
Ripple WJE, Larsen EJ, Renkin RA, Smith DW. 2001. Trophic cascades among wolves, elk and aspen on Yellowstone National Park's northern range. Biological Conservation. 102: 227-234.
Rothman RJ, Mech LD. 1979.Scent-marking in lone wolves in and newly formed pairs. Animal Behavior 27:750-760.
Sablin MV, Khlopachev GA. 2002. The earliest Ice Age dogs: evidence from Eliseevichi. Current Anthropology. 43(5): 795-799.
Savolainen P, Zhang YP, Luo J, Lundeberg J, Leitner T. 2002. Genetic evidence for an East Asian origin of domestic dogs. Science. 298: 1610-1613.
Schenkel R. 1947. Expression studies in wolves. Behaviour. 1(2): 81-129.
Schenkel R. 1967. Submission: Its features and function in the wolf and dog. American Zoologist. 7: 319-329.
Schonberner D. 1965. Observations on the reproductive biology of the wolf. Zeitschrift fur Saugertierkunde. 30(3): 171-178.
Seal US, Plotka ED, Packard JM, Mech LD. 1979. Endocrine correlates of reproduction in the wolf. I. Serum progesterone, estradiol and LH during the estrous cycle. Biology of Reproduction. 21: 1057-1066.
Shrestha TK. 1997. Mammals of Nepal: with references to those of India, Bangladesh, Bhutan, and Pakistan. Bimala Shrestha.
Siminski P. 2012. Mexican wolf, Canis lupus baileyi, International Studbook. Palm Desert (CA): The Living Desert.
Siminski P. 2013. Mexican wolf, Canis lupus baileyi, International Studbook. Annual Update [2013 Jul 31]. Palm Desert (CA): The Living Desert.
Siminski P, Spevak E, editors. 2013. Population analysis and breeding and transfer plan: Mexican wolf (Canis lupus baileyi) AZA species survival plan, yellow program. The Living Desert, Palm Desert (California).
Sullivan JO. 1978. Variability in the wolf, a group hunter. In: Hall RL, Sharp HS, eds. Wolf and man: evolution in parallel. New York: Academic Press.
Tedford RH, Wang X, et al. 2009. Phylogenetic systematics of the North American fossil Caninae (Carnivora, Canidae). Bulletin of the American Museum of Natural History. No. 325. Available from: http://digitallibrary.amnh.org/dspace/handle/2246/5999
Thalmann O, Shapiro B, Cui P, Schuenemann VJ, Sawyer SK, Greenfield DL, Germonpre MB, Sablin MB, Lopez-Giraldez F, Domingo-Roura X, Napierala H, Uerpmann H-P, Loponte DM, Acosta AA, Giemsch L, Schmitz RW, Worthington B, Buikstra JE, Bruzhkova A, Graphodatsky AS, Ovodov ND, Wahlberg N, Freedman AH, Schweizer RM, Koepfli K-P, Leonard JA, Meyer M, Krause J, Paabo S, Green RE, Wayne RK. 2013. Complete mitochondrial genomes of ancient canids suggest a European origin of domestic dogs. Science. 342: 871-874.
UNEP. 2019. Species+: Canis lupus [online database]. Nairobi, Kenya: UNEP [compiled by UNEP-WCMC, Cambridge, UK]. [accessed 2019 Sep 27]. https://www.speciesplus.net/#/taxon_concepts/4442/legal.
USFWS. 2019. Mexican wolf recovery program monthly update: August 1-31, 2019. Albuquerque (NM): U.S. Fish & Wildlife Service Southwest Region. https://www.fws.gov/southwest/es/mexicanwolf/recoverydocuments.html.
USFWS website. U.S. Fish and Wildlife Service, U.S. Department of the Interior. [cited 2014 Mar 01]. Available from: http://www.fws.gov/mountain-prairie/species/mammals/wolf/.
U.S. Fish & Wildlife Service (Midwest Region). 2019. Wolf — Western Great Lakes [Internet]. Bloomington (MN): U.S. Fish & Wildlife Service; [last updated 2017 Apr; accessed 2019 Apr 02]. https://www.fws.gov/midwest/wolf/.
vonHoldt BM, Pollinger JP, Lohmueller KE, Han E, Parker HG, et al. 2010. Genome-wide SNP and haplotype analyses reveal a rich history underlying dog domestication. Nature. 464: 898–902.
Wang D-d, Zhai W, Yang H-c, Fan R-x, Cao X, Zhong L, Wang L, Liu F, Wu H, Cheng L-g, Poyarkov AD, Poyarkov NA, Tang S-s, Zhao W-m, Gao Y, Lv X-m, Irwin DM, Savolainen P, Wu C-I, Zhang Y-p. 2013. The genomics of selection in dogs and the parallel evolution between dogs and humans. Nature Communications. 4: 1860.
Wayne RK. 1993. Molecular evolution of the dog family. Trends in Genetics. 9: 218-224.
Wydeven A. 2011. The history of wolves in Wisconsin. Madison (WI): State of Wisconsin, Department of Natural Resources.
Zeder MA. 2012. Pathways to animal domestication. In: Gepts P, et al, editors. Biodiversity in agriculture: domestication, evolution and sustainability. Cambridge: Cambridge University Press. Pp 227-259.
Zhang H, Wei Q, Zhang H, Chen L. 2011. Comparison of the fraction of olfactory receptor pseudogenes in wolf (Canis lupus) with domestic dog (Canis familiaris). Journal of Forestry Research. 22(2): 275-280.
Zimen E. 1976. On the regulation of pack size in wolves. Zeitschrift fur Tierpsychologie. Journal of Comparative Ecology. 40: 300-341.
